optimus prime is a character in the Transformers universe. He is the leader of the Autobots, a faction of heroic Transformers from the planet Cybertron who wage their battles against the evil forces of his brother's group the Decepticons for control of their homeworld, and by extension, peace in the universe. optimus-prime is depicted as a heroic, brave and compassionate character who puts all his talent to use to improve the world around him. Optimus has a strong sense of justice and righteousness, and has dedicated himself to the protection of all life, particularly the inhabitants of Earth; he will battle his foes with unyielding resolve to uphold this belief. [Wikipedia]
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The original optimus prime transforms into a cab over semi truck. The truck's cab transforms into the robot mode of Prime himself, with vast strength and armed with a laser rifle. Contained within his chest is the mystic talisman known as the Autobot Matrix of Leadership (or Creation Matrix, depending on the continuity), carried by all Autobot leaders and passed down through the ages. The comic book continuities tend to claim that Optimus' direct predecessor as Prime and thus bearer of the Matrix was Sentinel Prime, a fact which is part of the Dreamwave comics canon. In the cartoon continuity, it is unknown who Optimus inherited the talisman from, as he is already named as a Prime when Alpha Trion rebuilds him from his former self, which appears to indicate that Alpha Trion may have given Prime the Matrix at that point in time just after the transformation of Orion Pax into optimus-prime. [Wikipedia]
This change is depicted in the episode War Dawn. The truck's trailer disconnects and transforms into the Combat Deck, a mobile battle-station/command headquarters with an "Auto-Launcher" robot armed with assorted artillery and beam weapons. It can also serve as a radio antenna for battlefield communications between the Autobots. The Combat Deck can launch Prime's third component, Roller, a mobile scout buggy that can easily slip behind enemy lines. Optimus' senses are tied into Roller's, and can perceive what Roller does. Injury to one component is felt by each of the others, and while Prime could survive the destruction of either, despite the slight degree of autonomy they possess, the Combat Deck and Roller would not be able to survive without him. [Wikipedia]
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